Colombia, between the Meta, Casanare, and Capanaparo rivers, western part of the Department of Arauca: Resguardo Los Iguanitos, Municipality of Tame, Lower Cusay, to the north of the city of Tame; northwestern part of the Department of Casanare: Resguardo Caño Mochuelo-Hato Corozal, municipalities of Hato Corozal and Paz de Ariporo, between – to the north – the city of Tame (Department of Arauca) and – to the south – Paz de Ariporo; north-northwestern part of Vichada: Resguardo Santa Teresita del Tuparro, Municipality of Cumaribo, to the northwest of the centre of the Municipality of Cumaribo, between the rivers Tomo and Vichada, about 200km to the west of the Orinoco River; Resguardo La Pascua, municipalities of Puerto Carreño and Guacacías. Venezuela, Apure State, along the Upper Capanaparo River and its tributary Riecito.

Arauca Department, Cravo Norte municipality; Casanare Department, Capanaparo rivers and tributaries, El Merey, Esmeralda, Santa María, Betania, Mochuelo, San José de Ariporo; also in Vichada Department along south bank of the Meta River.

In Colombia: Arauca department: Cravo Norte municipality; Casanare department: Betania, El Merey, Esmeralda, Mochuelo, San José de Ariporo, and Santa María on Capanaparo River and tributaries; Vichada department: Meta River south bank.

In Venezuela: Apure state: upper Capanaparo River area and Riecito tributary.